
Irritator challengeri
Pronounced IR-ih-tay-tor chal-en-JER-eye
Tetanurae · Spinosauridae
A spinosaurid from the Early Cretaceous of north-east Brazil, known from the most complete spinosaurid skull ever found. Its long, narrow snout and conical teeth suited catching fish and other small prey on a tropical coastal lagoon.
- Bipedal
- Crested
- Possibly semiaquatic
- Period
- Cretaceous· Aptian
- Diet
- Piscivore
- Length
- 6–8 m
- Mass
- 1,000 kg
- Location
- Ceará, Brazil, South America
Quick answers
- How big was Irritator challengeri?
- Irritator challengeri was about 6–8 m long. It stood about 1.8–2.4 m tall at the hips. It weighed about 1 t. (Wikipedia: Irritator 2026; Paul 2016)
- When did Irritator challengeri live?
- Irritator challengeri lived during the Early Cretaceous, about 119.6–113.2 million years ago. (Paleobiology Database: Irritator challengeri … 2026; Schade et al. 2023)
- Where have Irritator challengeri fossils been found?
- Fossils of Irritator challengeri have been found in Romualdo Formation (Ceará) and Brazil. (Martill et al. 1996; Sues et al. 2002)
- What did Irritator challengeri eat?
- Irritator challengeri was a piscivore or carnivore. (Martill et al. 1996; Sues et al. 2002)
- Who named Irritator challengeri, and when?
- Irritator challengeri was described by David M. Martill et al. in 1996. (Martill et al. 1996; Schade et al. 2023)
On this page
IR-ih-tay-tor chal-en-JER-eye
- Period
- Cretaceous119.6–113.2 Ma
- Diet
- Piscivore
- Length
- 6–8 m
- Mass
- 1 t
- Location
- Brazil, South America
- Classification
- Tetanurae · Spinosauridae
- Described
- 1996, David M. Martill et al.
Overview
Irritator challengeri lived in the Araripe Basin of north-east Brazil in the late Aptian, a little over 113 million years ago. Its skull came out of a limestone concretion near Santana do Cariri, Ceará, and was sold by fossil dealers to the State Museum of Natural History in Stuttgart, Germany.
Dealers had built up the snout with plaster to make it look longer, which "irritated" the researchers who had to remove it; the species name honours Professor Challenger of Arthur Conan Doyle's "The Lost World". Martill and colleagues (1996) first thought it was a maniraptoran, and Sues and colleagues (2002) showed it was a spinosaurid.
CT scans by Schade and colleagues (2023) revealed almost the whole palate and braincase. The skull had binocular vision in front of the snout and a relatively weak but fast bite, and the lower jaws spread sideways when opening, like those of pelicans. It probably snapped up fish and small vertebrates, and a spinosaurid tooth has even been found in a pterosaur neck bone from the same rocks.
Name meaning
From "irritation", for the frustration caused by the dealers' alterations to the skull; challengeri after Professor Challenger in Conan Doyle's "The Lost World" (Martill et al. 1996).
Biology & evidence
What scientists think about how this animal lived, and how strong the evidence for each idea is.
Feeding
Irritator ate pterosaurs as well as fish.
Evidence level: HypothesisA spinosaurid tooth stuck in a pterosaur bone shows that some spinosaurid of the formation fed on one, by hunting or scavenging; it is not certain the tooth belonged to Irritator.
Evidence for / against · 1
Evidence for
Trace fossil · Spinosaurid tooth in a pterosaur vertebra (Romualdo Formation)
A spinosaurid tooth embedded in a pterosaur neck vertebra from the same formation.
Irritator had a relatively weak but fast bite suited to snapping small prey.
Evidence level: Moderate evidenceJaw mechanics and the inner ear suggest quick snaps at fish and small vertebrates rather than a crushing bite.
Evidence for / against · 2
Evidence for
Specimen · CT reconstruction of the Irritator skull
Medical and micro-CT reconstructions of SMNS 58022 reveal the palate and braincase, binocular vision in front of the snout, a weak but fast bite and lower jaws that spread sideways when opening.
Specimen: SMNS 58022
Comparative anatomy · Irritator braincase endocast
A digital endocast and inner ear indicating fast, well-controlled downward head movements and a snout held angled about 45 degrees down.
Specimen: SMNS 58022
Other
The Irritator holotype is the most complete spinosaurid skull known.
Evidence level: Direct evidenceOnly the tip of the snout and the front of the lower jaws are missing, while other spinosaurids are known mostly from snouts and isolated bones.
Evidence for / against · 1
Evidence for
Specimen · CT reconstruction of the Irritator skull
Medical and micro-CT reconstructions of SMNS 58022 reveal the palate and braincase, binocular vision in front of the snout, a weak but fast bite and lower jaws that spread sideways when opening.
Specimen: SMNS 58022
Size
- Length
- 6–8 m
- Height
- 2 mpublished range 1.8–2.4 m
- Mass
- 1,000 kg
(26.2 ft)
Based on SMNS 58022
(6.6 ft)
Timeline
Periods: Aptian (121.4–113 Ma), Early Cretaceous (145–100.5 Ma)
Late Aptian (Schade et al. 2023, after Arai & Assine 2020); PBDB gives the same bin, 119.57-113.2 Ma. Older works, and Wikipedia, date the Romualdo Formation to the Albian.
Location
Formations
- Romualdo Formation · Ceará
Places
- Near Buxexé, Santana do Cariri, Ceará · Romualdo Formation · Ceará · position approximate
These are the places where fossils have been found, not the full area where the animal lived.
Ecosystem
Lived in: Romualdo (Araripe Basin)
119.57–113.2 million years ago · marine margin, lake
A tropical coastal lagoon with irregular freshwater influence and marine incursions, surrounded by arid to semi-arid land; famous for fish, pterosaurs and soft tissues preserved in limestone concretions.
Classification
- cladeDinosauria
- cladeSaurischia
- cladeTheropoda
- cladeTetanurae
- superfamilyMegalosauroidea
- familySpinosauridae
- subfamilySpinosaurinae
- genusIrritator
- speciesIrritator challengeri
Taxonomic status: Valid
Fossil record
| Catalog number | Status | Institution | Found |
|---|---|---|---|
| SMNS 58022 | Holotype | Staatliches Museum für Naturkunde Stuttgart (SMNS) | — |
What has been found
Holotype: SMNS 58022
- Preserved
- Partial
- Reconstructed
- Unknown / not recorded
Region by region (table)
| Region | State |
|---|---|
| Skull | Partial · Missing most of the premaxillae and the front of the maxillae. |
| Mandible | Partial · Missing the front ends. |
| Teeth | Partial |
| Cervical vertebrae | Unknown / not recorded |
| Dorsal vertebrae | Unknown / not recorded |
| Sacrum | Unknown / not recorded |
| Caudal vertebrae | Unknown / not recorded |
| Ribs | Unknown / not recorded |
| Gastralia | Unknown / not recorded |
| Scapula / shoulder girdle | Unknown / not recorded |
| Humerus | Unknown / not recorded |
| Forearm (radius / ulna) | Unknown / not recorded |
| Hand (manus) | Unknown / not recorded |
| Pelvis | Unknown / not recorded |
| Femur | Unknown / not recorded |
| Tibia / fibula | Unknown / not recorded |
| Foot (pes) | Unknown / not recorded |
Gallery
Discovery
- Year described
- 1996
- Described by
- David M. Martill et al.
- Type specimen
- SMNS 58022
Discovery timeline
1996 · Formally described
Martill and colleagues name Irritator
Named as a crested maniraptoran, the first dinosaur named from the formation.
People: David M. Martill
2002 · Revision
Irritator redescribed as a spinosaurid
After further preparation, Sues and colleagues confirm it is a spinosaurid.
People: Hans-Dieter Sues, David M. Martill
2023 · Major study
CT reappraisal of the Irritator skull
Schade and colleagues reconstruct the whole skull digitally.
People: Marco Schade
Sources
- (2026). Paleobiology Database: Irritator challengeri occurrence and type collection 28052. The Paleobiology Database (paleobiodb.org), data service 1.2, accessed 2026-09-23. https://paleobiodb.org/data1.2/colls/list.json?coll_id=28052&show=paleoloc,strat,loc,ref,geo&pgm=gplates&vocab=pbdb (opens in a new tab)
- (2026). Wikipedia: Irritator. Wikipedia (English), accessed 2026-09-24. https://en.wikipedia.org/wiki/Irritator (opens in a new tab)
- Martill, D. M., Cruickshank, A. R. I., Frey, E., Small, P. G. & Clarke, M. (1996). A new crested maniraptoran dinosaur from the Santana Formation (Lower Cretaceous) of Brazil. Journal of the Geological Society 153: 5-8. doi:10.1144/gsjgs.153.1.0005 (opens in a new tab)
- Paul, G. S. (2016). The Princeton Field Guide to Dinosaurs (2nd edition). Princeton University Press.
- Schade, M., Rauhut, O. W. M. & Evers, S. W. (2020). Neuroanatomy of the spinosaurid Irritator challengeri (Dinosauria: Theropoda) indicates potential adaptations for piscivory. Scientific Reports 10: 9259. doi:10.1038/s41598-020-66261-w (opens in a new tab)
- Schade, M., Rauhut, O. W. M., Foth, C., Moleman, O. & Evers, S. W. (2023). A reappraisal of the cranial and mandibular osteology of the spinosaurid Irritator challengeri (Dinosauria: Theropoda). Palaeontologia Electronica 26(2): a17. doi:10.26879/1242 (opens in a new tab)
- Sues, H.-D., Frey, E., Martill, D. M. & Scott, D. M. (2002). Irritator challengeri, a spinosaurid (Dinosauria: Theropoda) from the Lower Cretaceous of Brazil. Journal of Vertebrate Paleontology 22: 535-547. doi:10.1671/0272-4634(2002)022[0535:ICASDT]2.0.CO;2 (opens in a new tab)