
Archaeopteryx lithographica
读音: AR-kee-OP-ter-iks lith-oh-GRAF-ih-kah
Avialae · Archaeopterygidae
A raven-sized feathered theropod from the Late Jurassic limestones of Solnhofen, Bavaria. Usually placed at the base of Avialae, the bird lineage, but it is a transitional animal, not the ancestor of modern birds.
- Feathered
- Bipedal
- Exceptional preservation
- Famous specimen
- Transitional fossil
- 时代
- 侏罗纪· 提塘阶
- 食性
- 肉食性
- 体长
- 0.5 m
- 体重
- 未知
- 地点
- Germany, Bavaria
快速问答
- Archaeopteryx lithographica有多大?
- Archaeopteryx lithographica体长约0.5 m。髋部高约0.1–0.2 m。 (Wellnhofer 2009; Paul 2016)
- Archaeopteryx lithographica生活在什么时候?
- Archaeopteryx lithographica生活在晚侏罗世,约1.492亿至1.45亿年前。 (Cohen et al. 2023; Rauhut et al. 2018)
- 在哪里发现过Archaeopteryx lithographica的化石?
- Archaeopteryx lithographica的化石发现于Solnhofen Formation (Bavaria)和Germany。 (von Meyer 1861; Xu et al. 2011)
- Archaeopteryx lithographica吃什么?
- Archaeopteryx lithographica是肉食性。 (von Meyer 1861; Xu et al. 2011)
- Archaeopteryx lithographica由谁命名,何时命名?
- Archaeopteryx lithographica由Hermann von Meyer于1861年描述。 (von Meyer 1861)
本页内容
- 时代
- 侏罗纪149.2–145 Ma
- 食性
- 肉食性
- 体长
- 0.5 m
- 体重
- 未知
- 产地
- Germany
- 分类
- Avialae · Archaeopterygidae
- 命名
- 1861, Hermann von Meyer
概览
此内容目前仅提供英文版。
Archaeopteryx lithographica lived about 150 million years ago, in the Late Jurassic, on islands in a warm lagoon sea that covered what is now southern Germany. Its fossils come from the fine lithographic limestones around Solnhofen and Eichstätt in Bavaria. Hermann von Meyer named it in 1861 from a single feather; a nearly complete skeleton, now in London, turned up the same year.
It mixed features of birds and of other small theropods: flight feathers on arms and tail, but also teeth, clawed fingers and a long bony tail. Found two years after On the Origin of Species, it became the classic example of a transitional fossil. Later specimens, such as the one described by Foth, Tischlinger and Rauhut (2014), show pennaceous feathers on the legs as well.
It is popularly called the "first bird", but that phrase needs care. Most analyses place it at or near the base of Avialae, the group that includes modern birds, yet Xu et al. (2011) recovered it closer to Deinonychus and Velociraptor. It is best read as an early member of the bird lineage, not the direct ancestor of living birds, and many feathered dinosaurs from China now blur the line it once drew.
名称含义
From Greek archaios ("ancient") and pteryx ("feather, wing"); lithographica refers to the lithographic limestone it was found in (von Meyer 1861).
生物学与证据
科学家对这种动物生活方式的看法,以及每种观点的证据有多充分。
运动方式
Archaeopteryx could fly actively, probably in short bursts.
证据等级:中等证据Its feathers have the asymmetric shape of flight feathers and its wing bones match those of birds that flap in short bursts. Its shoulder was less advanced than modern birds', so it was not a strong flier.
支持 / 反对证据 · 2
支持证据
组织学 · Archaeopteryx wing bone cross-sections
Synchrotron scans of wing bones show thin walls and a geometry closer to birds that flap in short bursts than to non-flying animals.
羽毛印痕 · Archaeopteryx flight feather impressions
Asymmetric flight feathers on the wings and a pair of rows along the tail, preserved in the limestone around the skeletons.
标本: NHMUK PV OR 37001 “London specimen”, MB.Av.101 “Berlin specimen”
皮肤与体表覆盖
At least some of the feathers of Archaeopteryx were black.
证据等级:中等证据Pigment structures preserved in one isolated feather match those of black feathers in living birds. It says nothing about the rest of the plumage, and the feather's owner is assumed to be Archaeopteryx.
支持 / 反对证据 · 1
支持证据
羽毛印痕 · Melanosomes in the isolated Archaeopteryx feather
Fossil pigment structures (melanosomes) in the 1861 feather, compared with living birds, indicate a dark, probably matte black colour.
羽毛
Archaeopteryx had flight feathers on its arms, tail and legs.
证据等级:直接证据The feathers are preserved as impressions in the fine limestone, including asymmetric wing feathers like those of flying birds and, in one specimen, pennaceous leg feathers.
支持 / 反对证据 · 1
支持证据
羽毛印痕 · Archaeopteryx flight feather impressions
Asymmetric flight feathers on the wings and a pair of rows along the tail, preserved in the limestone around the skeletons.
标本: NHMUK PV OR 37001 “London specimen”, MB.Av.101 “Berlin specimen”
其他
Archaeopteryx is an early member of the bird lineage (Avialae).
证据等级:中等证据Most family-tree analyses put it at the base of the bird lineage, but some place it closer to Velociraptor. It is best read as a close relative near the origin of birds, not the ancestor of living birds.
支持 / 反对证据 · 1
支持证据
系统发育推断 · Phylogenetic analyses placing Archaeopteryx
Most analyses recover Archaeopteryx as the earliest avialan; Xu et al. (2011) recovered it among deinonychosaurs instead.
体型
- 体长
- 0.5 m
- 身高
- 0.2 m文献范围 0.1–0.2 m
- 体重
- 未知
(1.6 英尺)
依据 Largest known specimens
(0.5 英尺)
依据 Berlin and Thermopolis specimens
时间线
时代: 提塘阶 (149.2–145 Ma), 晚侏罗世 (161.5–145 Ma)
Ages are the ICS 2023/09 bounds of the Tithonian, the stage(s) the fossils are assigned to; they bracket the record and are not dated first/last appearances. Rauhut et al. (2018) place the oldest specimen (Schamhaupten) at the Kimmeridgian/Tithonian boundary; most specimens are early Tithonian.
产地
地层
- Solnhofen Formation · Bavaria
- Langenaltheim near Solnhofen, Bavaria · Solnhofen Formation · Bavaria · 位置为近似值
这些是发现化石的地点,而不是该动物生活的全部范围。
生态系统
生活于: Solnhofen archipelago
152–150 百万年前 · 岛屿, 海洋边缘, 其他
Warm, shallow lagoons behind reefs, dotted with low, dry islands, on the northern margin of the Tethys Sea in the Late Jurassic. The stagnant, salty bottom water let fine lime mud bury carcasses in exceptional detail.
分类
- 演化支Dinosauria
- 演化支Saurischia
- 演化支Theropoda
- 演化支Coelurosauria
- 演化支Maniraptora
- 演化支Avialae
- 科Archaeopterygidae
- 属Archaeopteryx
- 种Archaeopteryx lithographica
分类地位: 有效
化石记录
| 编号 | 类型 | 收藏机构 | 发现年份 |
|---|---|---|---|
| NHMUK PV OR 37001 “London specimen” | 新模标本 | Natural History Museum, London (NHMUK) | 1861 |
| MB.Av.101 “Berlin specimen” | 归入标本 | Museum für Naturkunde (MB) | 1876 |
| WDC-CSG-100 “Thermopolis specimen” | 归入标本 | Wyoming Dinosaur Center (WDC) | — |
已发现的部分
新模标本: NHMUK PV OR 37001 “London specimen”
- 保存
- 部分保存
- 复原
- 未知 / 未记录
新模标本: NHMUK PV OR 37001 “London specimen”
按部位查看(表格)
| 部位 | 状态 |
|---|---|
| Skull | 部分保存 · Braincase only. |
| Mandible | 未知 / 未记录 |
| Teeth | 未知 / 未记录 |
| Cervical vertebrae | 未知 / 未记录 |
| Dorsal vertebrae | 保存 |
| Sacrum | 未知 / 未记录 |
| Caudal vertebrae | 保存 · Long bony tail with feathers. |
| Ribs | 未知 / 未记录 |
| Gastralia | 未知 / 未记录 |
| Scapula / shoulder girdle | 保存 · Furcula. |
| Humerus | 保存 |
| Forearm (radius / ulna) | 未知 / 未记录 |
| Hand (manus) | 保存 |
| Pelvis | 保存 |
| Femur | 保存 |
| Tibia / fibula | 未知 / 未记录 |
| Foot (pes) | 保存 |
归入标本: MB.Av.101 “Berlin specimen”
按部位查看(表格)
| 部位 | 状态 |
|---|---|
| Skull | 保存 |
| Mandible | 保存 |
| Teeth | 保存 |
| Cervical vertebrae | 保存 |
| Dorsal vertebrae | 保存 |
| Sacrum | 未知 / 未记录 |
| Caudal vertebrae | 保存 |
| Ribs | 未知 / 未记录 |
| Gastralia | 未知 / 未记录 |
| Scapula / shoulder girdle | 保存 |
| Humerus | 保存 |
| Forearm (radius / ulna) | 保存 |
| Hand (manus) | 保存 |
| Pelvis | 保存 |
| Femur | 保存 |
| Tibia / fibula | 保存 |
| Foot (pes) | 保存 |
归入标本: WDC-CSG-100 “Thermopolis specimen”
按部位查看(表格)
| 部位 | 状态 |
|---|---|
| Skull | 保存 |
| Mandible | 未知 / 未记录 |
| Teeth | 未知 / 未记录 |
| Cervical vertebrae | 未知 / 未记录 |
| Dorsal vertebrae | 未知 / 未记录 |
| Sacrum | 未知 / 未记录 |
| Caudal vertebrae | 未知 / 未记录 |
| Ribs | 未知 / 未记录 |
| Gastralia | 未知 / 未记录 |
| Scapula / shoulder girdle | 未知 / 未记录 |
| Humerus | 未知 / 未记录 |
| Forearm (radius / ulna) | 未知 / 未记录 |
| Hand (manus) | 保存 |
| Pelvis | 未知 / 未记录 |
| Femur | 未知 / 未记录 |
| Tibia / fibula | 未知 / 未记录 |
| Foot (pes) | 保存 |
发现
- 命名年份
- 1861
- 命名者
- Hermann von Meyer
- 模式标本
- NHMUK PV OR 37001 (London specimen; neotype replacing the original isolated feather)
发现时间线
1861 · 标本出土
The London specimen is found
A nearly complete skeleton with feathers is found near Langenaltheim and sold to the British Museum in 1862.
1861 · 首次发现材料
The Archaeopteryx feather is named
Hermann von Meyer names Archaeopteryx lithographica from an isolated feather found near Solnhofen.
人物:Hermann von Meyer
1863 · 重要研究
Owen describes the London specimen
Richard Owen describes the skeleton and interprets it as a bird.
人物:Richard Owen
1876 · 标本出土
The Berlin specimen is found
The most complete specimen, with its skull, is found near Eichstätt.
2011 · 修订
London specimen fixed as neotype
The ICZN designates the London skeleton as the name-bearing type, replacing the isolated feather.
参考文献
- Cohen, K. M., Harper, D. A. T., Gibbard, P. L. & Car, N. (2023). International Chronostratigraphic Chart (v2023/09). International Commission on Stratigraphy. https://stratigraphy.org/ICSchart/ChronostratChart2023-09.pdf(在新标签页中打开)
- Foth, C., Tischlinger, H. & Rauhut, O. W. M. (2014). New specimen of Archaeopteryx provides insights into the evolution of pennaceous feathers. Nature 511: 79-82. doi:10.1038/nature13467(在新标签页中打开)
- Paul, G. S. (2016). The Princeton Field Guide to Dinosaurs (2nd edition). Princeton University Press.
- Rauhut, O. W. M., Foth, C. & Tischlinger, H. (2018). The oldest Archaeopteryx (Theropoda: Avialiae): a new specimen from the Kimmeridgian/Tithonian boundary of Schamhaupten, Bavaria. PeerJ 6: e4191. doi:10.7717/peerj.4191(在新标签页中打开)
- von Meyer, H. (1861). Archaeopteryx lithographica (Vogel-Feder) und Pterodactylus von Solenhofen. Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefakten-Kunde 1861: 678-679.
- Wellnhofer, P. (2009). Archaeopteryx: The Icon of Evolution. Verlag Dr. Friedrich Pfeil, München.
- Xu, X., You, H., Du, K. & Han, F. (2011). An Archaeopteryx-like theropod from China and the origin of Avialae. Nature 475: 465-470. doi:10.1038/nature10288(在新标签页中打开)